Please cite this article in press Divya D Shirke et al., Borago Officinalis Linn: A Comprehensive Study On Bioactive Compounds And Its Various Pharmacological Uses, Indo Am. J. P. Sci, 2024; 11 (09).

1) Río‐Celestino MD, Font R, de Haro‐Bailón A. Distribution of fatty acids in edible organs and seed fractions of borage (Borago officinalis L.). Journal of the Science of Food and Agriculture. 2008 Jan 30;88(2):248-55.
2) Mansion G, Selvi F, Guggisberg A, Conti E. Origin of Mediterranean insular endemics in the Boraginales: integrative evidence from molecular dating and ancestral area reconstruction. Journal of Biogeography. 2009 Jul;36(7):1282-96.
3) Torabi F, Majd A, Enteshari S. Effect of exogenous silicon on germination and seedling establishment in Borago officinalis L. Journal of Medicinal Plants Research. 2012 Mar 16;6(10):1896-901.
4) Colombo ML, Assisi F, Della Puppa T, Moro P, Sesana FM, Bissoli M, Borghini R, Perego S, Galasso G, Banfi E, Davanzo F. Most commonly plant exposures and intoxications from outdoor toxic plants. Journal of pharmaceutical sciences and research. 2010 Jul 1;2(7):417.
5) Bianco VV, Santamaria P, Elia A. Nutritional value and nitrate content in edible wild species used in southern Italy. InIII International Symposium Diversification of Vegetable Crops 467 1996 Sep 24 (pp. 71-90).
6) Singh H, Du J, Yi TH. Green and rapid synthesis of silver nanoparticles using Borago officinalis leaf extract: anticancer and antibacterial activities. Artificial cells, nanomedicine, and biotechnology. 2017 Oct 3;45(7):1310-6.
7) De Lisi A, Montesano V, Negro D, Sarli G, Blanco E, Sonnante G, Laghetti G. Genetic diversity in Borago officinalis germplasm as revealed by seed oils and AFLP polymorphism. Genetic resources and crop evolution. 2014 Apr;61:853-9.
8) Sonnante G, De Paolis A, Pignone D. Relationships among artichoke cultivars and some related wild taxa based on AFLP markers. Plant Genetic Resources. 2003 Aug;1(2-3):125-33.
9) Gorenflo A, Diekötter T, van Kleunen M, Wolters V, Jauker F. Contrasting pollination efficiency and effectiveness among flower visitors of Malva sylvestris, Borago officinalis and Onobrychis viciifolia. Journal of pollination Ecology. 2017 May 29;21.
10) Borowy A, Chwil M, Kapłan M. Biologically active compounds and antioxidant activity of borage (Borago officinalis L.) flowers and leaves. Acta Scientiarum Polonorum Hortorum Cultus. 2017 Oct 31;16(5):169-80.
11) LUIS‐ARTEAGA MA, GARCÍA‐ARENAL FE, RODRÍGUEZ‐CEREZO EM. Characterization of a strain of clover yellow vein potyvirus infecting borage (Borago officinalis L.) in Spain. Plant pathology. 1996 Feb;45(1):38-44.
12) Akbarpour V, ARUEI H, Nemati SH. Phytochemical and morphological attributes of borage (Borago officinalis) affected by salicylic acid as an enhancer. Notulae Scientia Biologicae. 2014 Jun 10;6(2):138-42.
13) Ghorbel S, Nabli MA. Pollen, pistil and their interrelations in Borago officinalis and Heliotropium europaeum (Boraginaceae). Grana. 1998 Jan 1;37(4):203-14.
14) Descamps C, Quinet M, Baijot A, Jacquemart AL. Temperature and water stress affect plant–pollinator interactions in Borago officinalis (Boraginaceae). Ecology and Evolution. 2018 Mar;8(6):3443-56.
15) ARUEI H, AZIZI AM, AKBARPOUR V. ENHANCEMENT OF PHYTOCHEMICAL AND MORPHOLOGICAL ATTRIBUTES OF BORAGO OFFICINALIS BY SALICYLIC ACID.
16) Peiretti PG, Palmegiano GB, Salamano G. QUALITY AND FATTY ACID CONTENT OF BORAGE (BORAGO OFFICINALIS L.) DURING THE GROWTH CYCLE. Italian journal of food science. 2004 Apr 1;16(2).
17) Abu-Qaoud H, Shawarb N, Hussen F, Jaradat N, Shtaya M. Comparison of qualitative, quantitative analysis and antioxidant potential between wild and cultivated Borago officinalis leaves from palestine. Pakistan journal of pharmaceutical sciences. 2018 May 1;31(3).
18) Kareem AT, Hamad MN. Separation and Identification of Phenolic Acid from Borago officinalis (F: Boraginaceae) Cultivated in Iraq. Iraqi Journal of Pharmaceutical Sciences (P-ISSN 1683-3597 E-ISSN 2521-3512). 2020 Dec 29;29(2):139-51.
19) Wang H, Provan GJ, Helliwell K. Determination of rosmarinic acid and caffeic acid in aromatic herbs by HPLC. Food Chemistry. 2004 Sep 1;87(2):307-11.
20) Griffiths G, Brechany EY, Jackson FM, Christie WW, Stymne S, Stobart AK. Distribution and biosynthesis of stearidonic acid in leaves of Borago officinalis. Phytochemistry. 1996 Sep 1;43(2):381-6.
21) Sayanova O, Shewry PR, Napier JA. Characterization and expression of a fatty acid desaturase from Borago officinalis. Journal of experimental botany. 1999 Mar 1;50(332):411-2.
22) Velasco L, Goffman FD. Chemotaxonomic significance of fatty acids and tocopherols in Boraginaceae. Phytochemistry. 1999 Oct 1;52(3):423-6.
23) Eskin NM. Borage and evening primrose oil. European Journal of Lipid Science and Technology. 2008 Jul;110(7):651-4.
24) Wettasinghe M, Shahidi F. Antioxidant and free radical-scavenging properties of ethanolic extracts of defatted borage (Borago officinalis L.) seeds. Food chemistry. 1999 Dec 1;67(4):399-414.
25) Wettasinghe M, Shahidi F, Amarowicz R, Abou-Zaid MM. Phenolic acids in defatted seeds of borage (Borago officinalis L.). Food chemistry. 2001 Oct 1;75(1):49-56.
26) Sánchez‐Escalante A, Djenane D, Torrescano G, Beltrán JA, Roncales P. Antioxidant action of borage, rosemary, oregano, and ascorbic acid in beef patties packaged in modified atmosphere. Journal of Food Science. 2003 Jan;68(1):339-44.
27) Mhamdi B, Wannes WA, Bourgou S, Marzouk B. Biochemical characterization of borage (Borago officinalis L.) seeds. Journal of food biochemistry. 2009 Jun;33(3):331-41.
28) Amarowicz R, Kozlowska H, Shimoyamada M, Okubo K. Chromatographic analysis of rapeseed glucoside fractions. Polish journal of food and nutrition sciences. 1992;1(1).
29) Zemmouri H, Ammar S, Boumendjel A, Messarah M, El Feki A, Bouaziz M. Chemical composition and antioxidant activity of Borago officinalis L. leaf extract growing in Algeria. Arabian journal of chemistry. 2019 Dec 1;12(8):1954-63.
30) Singh M, Kamal Y, Khan M, Parveen R, Ahmad S. In vitro antioxidant activity and HPTLC analysis of Borago officinalis Linn. Indian Journal of Pharmaceutical Education and Research. 2013 Oct 1;47(4):24-30.
31) Aitzetmüller K, Grönheim M. Separation of highly unsaturated triacylglycerols by reversed phase HPLC with short wavelength UV detection. Journal of High Resolution Chromatography. 1992 Apr;15(4):219-26.
32) Galwey NW, Shirlin AJ. Selection of borage (Borago officinalis) as a seed crop for pharmaceutical uses. Heredity. 1990 Oct;65(2):249-57.
33) Ramezani M, Amiri MS, Zibaee E, Boghrati Z, Ayati Z, Sahebkar A, Emami SA. A review on the phytochemistry, ethnobotanical uses and pharmacology of Borago species. Current Pharmaceutical Design. 2020 Jan 1;26(1):110-28.
34) Rabiei Z, Lorigooini Z, Kopaei MR. Effects of hydroalcoholic extract of Borago officinalis on naloxone-precipitated withdrawal syndrome in morphine-dependent mice. Bangladesh Journal of Pharmacology. 2016 Oct 1;11(4):824-9.
35) Ghahremanitamadon F, Shahidi S, Zargooshnia S, Nikkhah A, Ranjbar A, Soleimani Asl S. Protective effects of Borago officinalis extract on amyloid β-peptide (25–35)-induced memory impairment in male rats: a behavioral study. BioMed research international. 2014 Oct;2014.
36) Samy MN, Hamed AN, Sugimoto S, Otsuka H, Kamel MS, Matsunami K. Officinalioside, a new lignan glucoside from Borago officinalis L. Natural product research. 2016 Apr 17;30(8):967-72.
37) Manthena SS, Polimati H, Annam SS, Hieu HV. Hepatoprotective Properties of Starflowers from an Annual Herb, Borago officinalis L.(Boraginaceae). International Pharmacy Acta. 2022 Oct;5(1):e7-1.
38) Hamed AN, Wahid A. Hepatoprotective activity of Borago officinalis extract against CCl4-induced hepatotoxicity in rats. J. of Nat. Prod. 2015;8:113-22.
39) Gilani AH, Bashir S, Khan AU. Pharmacological basis for the use of Borago officinalis in gastrointestinal, respiratory and cardiovascular disorders. Journal of Ethnopharmacology. 2007 Dec 3;114(3):393-9.
40) Mhamdi B, Wannes WA, Sriti J, Jellali I, Ksouri R, Marzouk B. Effect of harvesting time on phenolic compounds and antiradical scavenging activity of Borago officinalis seed extracts. Industrial Crops and Products. 2010 Jan 1;31(1):e1-4.
41) Komaki A, Rasouli B, Shahidi S. Anxiolytic effect of Borago officinalis (Boraginaceae) extract in male rats. Avicenna Journal of Neuro Psycho Physiology. 2015 Feb 10;2(1):34-8.